Along with Metchnikoff, Russia's leading 19th century experimental biologist. A Darwinian whose careful studies of invertebrate development and efforts to find the nearest relatives of the vertebrates (such as Amphioxus) were crucial to late 19th century phylogenetic debates. Brother of the paleontologist Vladimir Kowalevsky. The modern Romanization of his name is Aleksandr Onufrievich Kovalevskii.
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