McGuire, J.L., J.J. Lawler, B.H. McRae, T. Nuñez, and D.M. Theobald. 2016. Achieving climate connectivity in a fragmented landscape. PNAS. doi: 10.1073/pnas.1602817113

McRae B.H., K. Popper, A. Jones, M. Schindel, S. Buttrick, K. Hall, R.S. Unnasch, and J.T. Platt. 2016. Conserving Nature’s Stage: Mapping Omnidirectional Connectivity for Resilient Terrestrial Landscapes in the Pacific Northwest. The Nature Conservancy, Portland Oregon. 47 pp. Available at:

McRae, B.H., V.B. Shah, and A. Edelman. 2016. Circuitscape: Modeling Landscape Connectivity to Promote Conservation and Human Health. The Nature Conservancy, Fort Collins, CO. 14 pp

Belote, R.T., M.S. Dietz, B.H. McRae, D.M. Theobald, M.L. McClure, G.H. Irwin, P.S. McKinley, J.A. Gage, and G.H. Aplet. 2016. Identifying Corridors among large protected areas in the United States. PLoS ONE 11(4): p.e0154223.

Heiner M, B.H. McRae, B. Lkhagvasuren, N. Batsaikhan, D. Galbadrakh, B. Tsogtsaikhan, B. Naranzul, O. Binderya, and J. Kiesecker. 2016. Modeling habitat connectivity of a nomadic migrant facing rapid infrastructure development:  Khulan habitat connectivity in the Southeast Gobi Region, Mongolia.  In: Capacity building for Mongolian Ministry of Environment and Green Development in relation to biodiversity and conservation in the southern Gobi Desert: Final Report. The Nature Conservancy Mongolia Program. Ulaanbaatar, Mongolia 

Krosby, M., I. Breckheimer, D. J. Pierce, P.H. Singleton, S.A. Hall, K.C. Halupka, W.L. Gaines, R.A. Long, B.H. McRae, B.L. Cosentino, and J.P. Schuett-Hames. 2015. Focal species and landscape “naturalness” corridor models offer complementary approaches for connectivity conservation planning. Landscape Ecology 30(10): 2121-2132.

Dutta, T., S. Sharma, P.S. Roy, B.H. McRae, and R. DeFries. 2015. Connecting the dots: Connectivity mapping for tigers in central India. Regional Environmental Change 1-15.

Nogeire, T., E. Boydston, K. Crooks, B.H. McRae, L. Lyren, and F. Davis. 2015. Can orchards help connect Mediterranean ecosystems? Animal movement data alter conservation priorities. American Midland Naturalist 174(1):105-116

Parks, L.C., D.O. Wallin, S.A. Cushman,  and B.H. McRae. 2015. Landscape-level analysis of mountain goat population connectivity in Washington and southern British Columbia. Conservation Genetics 16(5): 1195-1207

Cushman, S.A., B.H. McRae, and K. McGarigal. 2015. Basics of landscape ecology: an introduction to landscapes and population processes for landscape geneticists. Chapter 2 in Balkenhol, N., S.A. Cushman, A.T. Storfer, and L.P. Waits, eds. Landscape Genetics: Concepts, Methods, Applications. Wiley.

Spear, S.F., S.A. Cushman, and B.H. McRae. 2015. Resistance surface modeling in landscape genetics.  Chapter 8 in Balkenhol, N., S.A. Cushman, A.T. Storfer, and L.P. Waits, eds. Landscape Genetics: Concepts, Methods, Applications. Wiley.

Torrubia, S., B.H. McRae, J.J. Lawler, S.A Hall, M. Halabisky, J. Langdon, and M. Case. 2014. Getting the most connectivity per conservation dollar. Frontiers in Ecology and the Environment. 12(9): 491–497

Lawler, J.J., A.S. Ruesch, J.D. Olden, and B.H. McRae. 2013. Projected climate-driven faunal movement routes. Ecology Letters 16(8):1014-1022.

Dickson B.G., G.W. Roemer, B.H. McRae, and J.M. Rundall. 2013. Models of Regional Habitat Quality and Connectivity for Pumas (Puma concolor) in the Southwestern United States. PLoS ONE 8(12): e81898.

Nuñez, T., J.J. Lawler, B.H. McRae, D.J. Pierce, M.R. Krosby, D.M. Kavanagh, P.H. Singleton, and J. . Tewksbury. 2013. Connectivity planning to facilitate species movements in response to climate change. Conservation Biology 27: 407–416.

McRae, B.H., S.A Hall, P. Beier, and D.M. Theobald. 2012. Where to restore ecological connectivity? Detecting barriers and quantifying restoration benefits. PLoS ONE 7(12): e52604. doi:10.1371/journal.pone.0052604

Baldwin R, S.E. Reed, B.H. McRae, D.M. Theobald, and R.W. Sutherland. 2012. Connectivity restoration in large landscapes: modeling landscape condition and ecological flows. Ecological Restoration 30 (4) 274–279.

Carroll, C., B.H. McRae,  and A. Brookes. 2012. Use of linkage mapping and centrality analysis across habitat gradients to conserve connectivity of gray wolf populations in western North America. Conservation Biology, 26(1), 78-87.

Beier, P., W. Spencer, R. Baldwin, and B.H. McRae. 2011. Best science practices for developing regional connectivity maps. Conservation Biology 25(5): 879-892

Schloss, C. A., Lawler, J. J., Larson, E. R., Papendick, H. L., Case, M. J., Evans, D. M., DeLap, J. H., et al. (2011). Systematic conservation planning in the face of climate change: bet-hedging on the Columbia Plateau. PloS ONE, 6(12), e28788.

Krosby, M., J. Hoffman, J.J. Lawler, and B.H. McRae. 2012. Integrating climate change into conservation planning in Washington State and the Pacific Northwest. Chapter 10 In: C.C. Chester, J.A. Hilty, and M.S. Cross, eds. Conservation and Climate Disruption: Ecoregional Science and Practice in a Changing Climate. Island Press.

Washington Wildlife Habitat Connectivity Working Group (WHCWG). 2012. Washington Connected Landscapes Project: Columbia Plateau Analysis. Washington Departments of Fish and Wildlife, and Transportation, Olympia, WA. Available at:

Washington Wildlife Habitat Connectivity Working Group (WHCWG). 2011. Washington Connected Landscapes Project: Climate Gradient Corridor Analysis. Washington Departments of Fish and Wildlife, and Transportation, Olympia, WA. Available at:

Safner,T., M.P. Miller, B.H. McRae, M.-J. Fortin, and S. Manel. 2011.

Comparison of Bayesian clustering and edge detection methods for inferring

boundaries in landscape genetics. International Journal of Molecular Sciences

12(2): 865-889.

Spear, S.F., N. Balkenhol, M.-J. Fortin, B.H. McRae and K. Scribner. 2010. Use

of resistance surfaces for landscape genetic studies: Considerations of

parameterization and analysis. Molecular Ecology 19(17): 3576-3591.

Epperson B.K., McRae B.H., Scribner K, Cushman S.A., Rosenberg M.S., Fortin

M.J., James P.M., Murphy M, Manel S, Legendre P, Dale M.R. 2010. Utility of

computer simulations in landscape genetics. Utility of computer simulations in

landscape genetics. Molecular Ecology 19(17):3549-3564

Washington Wildlife Habitat Connectivity Working Group (WHCWG). 2010.

Washington Connected Landscapes Project: Statewide Analysis. Washington

Departments of Fish and Wildlife, and Transportation, Olympia, WA. Available


Lee-Yaw J.A., Davidson A., McRae B.H., Green D.M. 2009. Do landscape processes predict phylogeographic patterns in the wood frog? Molecular Ecology 18(9):1863-1874

McRae, B.H., B.G. Dickson, T.H. Keitt, and V.B. Shah. 2008. Using circuit theory to model connectivity in ecology and conservation. Ecology 10: 2712-2724.

Shah,V.B. and B.H. McRae. 2008. Circuitscape: a tool for landscape ecology. In: G. Varoquaux, T. Vaught, J. Millman (Eds.). Proceedings of the 7th Python in Science Conference (SciPy 2008), pp. 62-66.

McRae, B.H., N.H. Schumaker, R.B. McKane, R.T.Busing, A.M. Solomon, and C.A. Burdick. 2008. A multi-model framework for simulating wildlife population response to land-use and climate change. Ecological Modelling 19:77-91.

McRae, B.H. and P. Beier. 2007. Circuit theory predicts gene flow in plant and animal populations. Proceedings of the National Academy of Sciences of the USA 104:19885-19890.

McRae, B.H. 2006. Isolation by resistance. Evolution 60:1551-1561.

McRae, B.H., P. Beier, L.Y. Huynh, L. DeWald, and P. Keim. 2005. Habitat barriers limit gene flow and illuminate historical events in a wide ranging carnivore, the American puma. Molecular Ecology 14:1965-1977.

Loeser, M.R., B.H. McRae, M.M. Howe, and T.G. Whitham. 2006. Litter hovels as havens for riparian spiders in an unregulated river. Wetlands 26:13-19.

McRae, B.H. 2002. Review of Logan, K. and L. Sweanor. 2001. Desert Puma: Evolutionary ecology and conservation of an enduring carnivore. Island Press. In Journal of Mammalogy 83(3):913-915.

McKinney , T. and B.H. McRae. 2001. Genetic analysis of feces for estimating size of a Sonoran Desert mountain lion population. Proceedings of the 2001 Desert Bighorn Conference, April 2001, Hermosillo, Mexico.

Harrod, R.J., B.H. McRae, and W.E. Hartl. 1998. Historical stand reconstruction in ponderosa pine forests to guide silvicultural prescriptions. Forest Ecology and Management 4499:1-14.