1. Hartter, J.*, Ryan, S.J.*, Southworth, J., and Chapman, C.A. 2011. Landscapes as continuous entities: forest disturbance and recovery in the Albertine Rift Landscape. Landscape Ecology 26(6):877-890 [PDF]

  2. Ryan, S.J., 2011. Invited Book Review. "Governing Africa's Forests in a Globalized World Laura A German, Alain Karsenty and Anne-Marie Tiani (Editors)". Natural Resources Forum 35(2): 146-147. [PDF]

  3. Stampone, M., Hartter, J., Chapman, C.A. , Ryan, S.J.  2011. Localized precipitation trends in and around a forest park in east equatorial Africa. Research Journal of Environmental and Earth Sciences 3(1):14-23 [PDF]

  4. Petorelli, N., Ryan, S.J., Mueller, T., Bunnefeld, N., Jedrzejewska, B., Lima, M., Kausrud, K. 2011. The Normalized Difference Vegetation Index (NDVI) in ecology: a decade of unforseen successes. Climate Research 46:15-27 [PDF]

  5. Hartter, J. and Ryan, S.J. 2010. Top-down or bottom-up? Decentralization, natural resource management and usufruct rights in the forests and wetlands of Western Uganda. Land Use Policy 27(3): 815-826. [PDF]

  6. Ladau, J. and Ryan, S.J. 2010. MPowering ecologists: community assembly tools for community assembly rules. Oikos 119(7): p. 1064-1069 [PDF]

  7. Bar-David, S, Bar-David, I, Cross, P.C. Ryan, S.J., Knechtel, C.U., Getz, W.M. 2009. Methods for assessing movement path recursion with application to African buffalo in South Africa. Ecology 90(9): 2467–2479 [PDF].

  8. Sánchez, M.S., Lloyd-Smith, J.O., Williams, B.G., Porco, T.C., Ryan, S.J., Borgdorff, M.W., Mansoer, J., Dye, C. and Getz, W.M. 2009. Incongruent HIV and tuberculosis co-dynamics in Kenya: Interacting epidemics monitor each other. Epidemics 1(1):14-20 [PDF]

  9. Ryan, S.J., Starks, P.T., Milton, K. and Getz, W.M. 2008. Intersexual conflict and group size in Alouatta palliata: a 23-year evaluation.  Int. J. Primatology. 29:405-420  [PDF]

  10. Gusset, M.*& Ryan, S.J.*, Hofmeyr, M., Van Dyk, G., Davies-Mostert, H.T., Graf, J.A., Owen, C., Szykman, M., Macdonald, D.W., Monfort, S.L., Wildt, D.E., Maddock, A.H., Mills, M.G.L., Slotow, R., Somers, M.J. 2008. Efforts going to the dogs? Evaluating attempts to re-introduce endangered wild dogs in South Africa. Journal of Applied Ecology 45:100-108  [PDF]

  11. Ryan, S.J., Knechtel, C.U. and Getz, W.M. 2007. Ecological cues, gestation length and birth timing in African Buffalo (Syncerus caffer). Behavioral Ecology 18:635-644 [PDF]

  12. Getz, W.M., Fortmann-Roe, S, Cross, P.C., Lyons, A.J. Ryan, S.J., et al. 2007. LoCoH: Nonparametric kernel methods for constructing home ranges and utilization distributions. PLoS ONE 2(2) [PDF]

  13. Ryan, S.J. 2006. The role of culture in conservation of small or endangered populations. Journal of Conservation Biology 20(4):1321-1324 [PDF]

  14. Ryan, S.J., Knechtel, C.U. and Getz, W.M.2006. Range and habitat selection of African buffalo in South Africa. Journal of Wildlife Management 70(3): 764-776 [PDF]

  15. Redfern, J.V., Ryan, S.J. and Getz, W.M. 2006. Defining Herbivore Assemblages in the Kruger National Park: A Correlative Coherence Approach. Oecologia 146:632-640 [PDF]

  16. Ryan, S.J., and Getz, W.M. 2005. A spatial location allocation GIS framework for managing water sources in a savanna nature reserve. South African Journal of Wildlife Research 35(2):153-178 [PDF]

  17. Ryan, S.J. and Jordaan, W. 2005. Activity Patterns of African Buffalo Syncerus caffer in the Lower Sabie Region, Kruger National Park, South Africa. Koedoe 48(2): 117-124 [PDF]

  18. Faust, L.J., Thompson, S.D., Earnhardt, J.M., Brown, E., Ryan, S.J., Sherman, M. and Yurenka, M. 2003. Using Stage-Based System Dynamics Modeling for Demographic Management of Captive Populations. Zoo Biology, 22:1, Wiley-Liss [PDF]

  19. Ryan, S., Roth, A.M., Thompson,S.D. and Gold, K.C. 2002. Effects of hand rearing on the reproductive success of Western Lowland Gorillas in North America. Zoo Biology 21:3, Wiley-Liss [PDF]

  20. Ryan, S.J. and Thompson, S.D., 2001. Disease Risk and Inter-institutional Transfer of Specimens in Cooperative Breeding Programs: Herpes and the Elephant Species Survival Plans, Zoo Biology 20:2, Wiley-Liss [PDF]

*Featured in “Dog days for evidence” Journal Watch Online, Society for Conservation Biology, July 31, 2007


Ryan, S.J., Cross, P.C., Winnie, J, Jr, Hay, C., Bowers, J., Getz, W.M. in revision. The utility of NDVI as a proxy for forage quality in a savanna landscape.

O'Leary, J. and Ryan, S.J. in review. Managing for trophic integrity in terrestrial and marine protected areas: case studies from Africa.

Wolf, A., Anderegg, W., Ryan, S.J., Christensen, J. in review. Robust detection of plant species range shifts under biased sampling regimes.

Rudnick, D., Preziosi, D., Epps, C., Deiffenbach, F., Kintsch, J., Beier, P., Jenness, J., Perkl., R., Trombulak, S., Ryan, S.J., Hartter, J., Gerber, L., Merenlender, A., Austen, D., Cushman, S. in review. Emerging principles in understanding landscape connectivity: practical tools for conservation decision-making. Issues in Ecology.

Hartter, J., Stampone, M.D., Ryan, S.J., Kirner, K., Chapman, C.A., Goldman, A., in reivew. Patterns and perceptions of climate change around a biodiversity hotspot

Ryan, S.J., Brashares, J.S., Chapman, C.A., Milbers, K., Walsh, C., Kilroy, C. in review. Gastrointestinal parasites of olive baboons (Papio anubis) in human-use areas of Mole National Park, Ghana.

Tildesley, M.J. and Ryan, S.J. in prep. The utility of land cover maps to inform spatial epidemic models of disease transmission in the UK livestock industry.

Ryan, S.J., Dobson, A.P. and Jones, J.H. in prep. The effects of contact structure, demography and movement on disease transmission within a structured primate metapopulation.

  1. Hartter, J., Ryan, S.J., Southworth, J., and Chapman, C.A. 2010. "Fortresses and Fragments: Impacts of Fragmentation in a Forest Park Landscape". in Proceedings of IUFRO Landscape Ecology International Conference. Braganca, Portugal [PDF]

  2. Chapman, C.A., Huffman,M.A., Ryan,S.J., Sengupta, R., Goldberg, T. 2009. “Ways forward in the study of primate parasite ecology”. in Primate parasite ecology: the dynamics and study of host-parasite relationships. Huffman, M.A. and Chapman, C.A., eds. Cambridge Studies in Biological and Evolutionary Anthropology. Cambridge University Press. [PDF]

  3. Ryan, S., Roth, A.M., Thompson, S.D. and Gold, K.C. 2001. "Effects of hand rearing on the reproductive success of Western Lowland Gorillas in North America". in The Apes: Challenges for the 21st Century. Chicago, Chicago Zoological Society, p.99. [PDF]

  4. Dobson, A., Ralls. K., Foster, M., Soule, M.,Simberloff, D., Doak, D., Estes, J., Mills, S., Mattson, D., Dirzo, R., Arita, H., Ryan, S., Norse, E., Noss, R., Johns, D. 1999. "Corridors: Maintaining Flows in Fragmented Landscapes". In Continental Conservation, Michael E. Soule and John Terborgh, eds. The Wildlands Project,Island Press.


Ryan, S.J. 2006. Spatial Ecology of African Buffalo and their Resources in a Savanna Ecosystem. PhD Dissertation, Department of Environmental Science, Policy and Management, UC Berkeley. [PDF]

Ryan, S.J. 1998. Primate Concerns: Development of a Rapid Censusing Technique, Social Effects on Pathogen Transmission and an Examination of Primate Conservation. BA Thesis, Department of Ecology and Evolutionary Biology, Princeton University [PDF] *incomplete document*


Smith, C, with Ouzman, S., Ryan, S. and Schonfeldt-Aultman, S. 2003. Renboogtaal in Renboogdorp gepraat. Die Volksblad 15 May


Ryan, S.J., (ed.). 1999. North American Regional Studbook for Sunbittern (Eurypyga helias). AZA /Lincoln Park Zoo

Ryan, S.J., (ed.). 1999. North American Regional Studbook for Pygmy Hippos (Hexaprotodon liberiensis). AZA/LPZ

Ryan, S.J., (ed.). 2000. North American Regional Studbook for Snow Leopards (Pantera uncia). AZA /LPZ

Ryan, S.J., (ed.). 2000. AZA Research Coordinators Directory. AZA/LPZ

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